Sunflower Family

California Aster © KKorbholz

Asteraceae (ass-ter-AY-see-ee)

Iconic Features

    • Composite heads of many flowers
    • Disk and/or straplike flowers

    Description (Jepson)

      • Eudicotyledons (eudicots) – a major lineage of flowering plants including most plants traditionally described as dicots and generally characterized by
        • 2 seed leaves (dicotyledon)
        • Netted (reticulate) leaf venation
        • Flower parts in fours and fives
        • Pollen grains with 3 pores (tricolpate)
        • Vascular bundles in stem arranged in a ring
        • Taproot system
      • Mostly annuals and herbaceous perennials
        • Also can be shrubs, vines, and trees
      • Leaves
        • Simple (not divided into leaflets) to compound (divided into leaflets)
        • Basal and/or cauline (on the stem)
        • Alternate (1 leaf at each junction with stem) or opposite (2 leaves at each junction with stem) or, rarely, whorled (3 or more leaves at each junction with stem)
      • Flowers
        • Inflorescence (flower arrangement) is in densely packed heads (capitulum/a) of dozens to hundreds of individual flowers
          • Heads may appear singly or in a variety of secondary arrangements
        • Flowerheads come in 4 basic forms (with many variations)
          • Radiate Heads have both disk and ray flowers (e.g. mule’s ears)
            • 5-lobed star- or tube-shaped disk flowers cluster in the center
            • 3-lobed strap-like ray flowers rim the edge
          • Ligulate (Liguliflorous) Heads have only 5-lobed, strap-like flowers (e.g. dandelion)
          • Discoid Heads have only 5-lobed star- or tube-shaped disk flowers (e.g. mugwort)
          • Disciform Heads have disk and disk-like flowers (e.g. everlasting)
            • 5-lobed star- or tube-shaped disk flowers cluster in the center
            • Disk-like flowers with minute or missing rays rim the edge
        • Flowers are protandrous (male flower parts mature before female parts) and employ a strategy of secondary pollen presentation
          • Anthers (pollen-producing part of the stamen/male structure) are fused into a column facing toward and surrounding the pistil (female flower part)
          • When still immature, the style (pistil stalk) elongates, pushing ripe pollen up through the anther column to present to pollinators
          • When the flower’s own pollen is depleted, the style splits to receive pollen from other flowers
          • This strategy is believed to reduce the chance of self-pollination
        • Sepals (protective covering for bud) are absent or modified into a pappus, composed of bristles or scales (e.g. the tufts of a dandelion seed)
        • Ovary inferior (below the attachment of other flower parts)
        • Composite head is usually held in a whorl of bracts (modified leaves), called an involucre
          • Individual bract is called a phyllary
        • Individual flowers lack stems; instead they are attached to a platform called a receptacle
      • Fruit is an achene (a single-seeded, dry fruit that does not split open), often with pappus attached to aid dispersal
      Radiate Head (L), Ligulate Head (M), Discoid Head (R) © JMason

      Notes

        • Approximately 23,000 species worldwide, in many habitats
          • Largest family of vascular plants in California (Jepson eFlora) and of eudicots globally
          • Found everywhere except Antarctica and the extreme Arctic
        • Great variety of horticultural, economic, and medicinal uses
        • Scientific name from the included genus Aster, from the Greek for “star”
          • An early name for this family was Compositae because what appears to be an individual flower is actually a composite of flowers, as in a bouquet
          • Other common names are the Daisy or Composite family
        • Due to its size, the Sunflower family is traditionally broken into a number of subgroups, including tribes; here are some tribes and representative plants
          • Mayweed Tribe (Anthemideae) – yarrow (Achillea millefolium), sagebrush (Artemisia species), pineapple weed (Chamomilla sauveolens)
          • Aster Tribe (Astereae) – asters (Aster species), gumplants (Grindelia species), coyote brush (Baccharis pilularis)
          • Chicory Tribe (Cichorieae) – California dandelion (Agoseris grandiflora), hawkweeds (Hieracium species), silver puffs (Uropappus lindleyi)
          • Thistle Tribe (Cynareae) – star-thistles (Centaurea species), Italian thistle (Carduus pycnocephalus)
          • Sunflower Tribe (Heliantheae) – mule ears (Wyethia species), tarweeds (Madia species), goldfields (Lasthenia species)
          • Everlasting Tribe (Inuleae) – cudweed (Pseudognaphalium californicum), stinkwort (Dittrichia graveolens)
        • Represented by 91 species at Edgewood

        See General References

        Specific References

          An Intriguing Way of Presenting One’s Pollen. 2019, Sep. 22. In Defense of Plants.

            Mason, J. 2004. [Sunflower family flower-head illustrations, adapted]. In T. Corelli, Flowering Plants of Edgewood Natural Preserve, 2nd. ed. Monocot Press, Half Moon Bay, California. (c) CC BY NC 3.0.

            Browse Some Edgewood Plants in this Family